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Publications & Books


In press

  • Chaparro, O.R., L.A. Lincoqueo, A.J. Schmidt, D. Veliz and J.A. Pechenik. Oxygen constraints in the brood chamber: Anaerobic metabolism and the payment of oxygen debt in females and in brooded embryos of the limpet Crepipatella dilatata. Biological Bulletin


  • Averbuj, A., Buchner-Miranda, J.A., Salas-Yanquin, L.P., Navarro, J.M., Pardo, L.M., Matos, A.S., Pechenik, J.A., and O.R. Chaparro. Energetic trade-offs: Implications for selection between two bivalve prey species by a carnivorous muricid gastropod. PLoS One 16(4): e0250937. https://doi.org/10.1371/journal.pone.0250937
  • Montory, J. A., Chaparro, O.R., Salas-Yanquin, L.P., Büchner-Miranda, J.A., Pechenik, J.A., and V.M. Cubillos. 2021.  Impact of intertidal distribution on the physiological performance of the filter-feeder bivalve Perumytilus purpuratus (Bivalvia, Mytilidae) from Southern Chile: Malacologia, 64(1): 137-147 https://doi.org/10.4002/040.064.0108
  • Salas-Yanquin, L.P., J.A. Buchner-Miranda, J.A. Montory, J.A. Pechenik, V.M Cubillos, A.S. Matos, and O.R. Chaparro. 2021. Relationship between over-crowding within egg capsules of the marine gastropod Acanthina monodon and prospects for juvenile success.  Marine Environmental Research 169: 105353  https://doi.org/10.1016/j.marenvres.2021.105353


  • McVey, M. and J.A. Pechenik. 2020. Using poetry in the undergraduate biology classroom.  The American Biology Teacher 82:1–5. https://doi.org/10.1525/abt.2020.82.6.1
  • Pechenik, J.A.,Richards, K.B., Frieliech, C.A., and C.M. Diederich.
    Examining the impact of a symbiotic lifestyle on the fecundity of the marine gastropod Crepidula plana. Invertebrate Biology e:122904. https://doi.org/10.1111/ivb.12294
  • Chaparro, O.R., Salas-Yanquin, L.P., Büchner-Miranda, J.A., Pechenik, J.A., Gray, M.W., Navarro, J.M., and V.M. Cubillos. Respiratory and desiccation constraints during encapsulated intertidal development of the marine gastropod Acanthina monodon. Marine Environmental Research 161: 105120
  • Pechenik, J.A., O.R. Chaparro,  Z.M. Lazarus, G.V. Tellado, E.M. Ostapovich, and D. Clark. 2020. Impact of short-term elevated temperature stress on winter-acclimated individuals of the marine gastropod Crepidula fornicataMarine Environmental Research 162: 105180.   https://doi.org/10.1016/j.marenvres.2020.105180
  • Rivera-Figueroa, G.A.,  J. A. Büchner-Miranda, L. P. Salas-Yanquin, J. A. Montory, V. M. Cubillos, J. A. Pechenik and O. R. Chaparro. 2021. Capture of conspecific planktonic larvae by the suspension-feeding gastropod Crepipatella peruviana: association between adult and larval size.  Journal of the Malacological Society of London, Vol. 87.  https://doi.org/10.1093/mollus/eyab002


  • Bogan S.N., J.B. McMahon, J.A. Pechenik, & A. Pires.  2019. Legacy of multiple stressors: responses of gastropod larvae and juveniles to ocean acidification and nutrition.  Biological Bulletin 236: 159-173.
  • Pechenik, J.A. O.R. Chaparro, A. Franklin, M.L. Mardones, & J.A. Montory. 2019.
    Thermal tolerance of intertidal and subtidal adults and embryos of the marine gastropod Crepipatella peruviana. Marine Ecology Progress Series 616: 67-81.
  • Pechenik, J.A., M. Levy, and J.D. Allen. 2019. Instant Ocean versus natural seawater: Impacts on aspects of reproduction and development in three marine invertebrates.  Biological Bulletin 237: 16-25. (Selected as “Editor’s Choice” for this issue)
  • Pechenik, J.A., A. Pires, J. Trudel, M. Levy T. Dooley, A. Resnikoff, and R. E. Taylor. 2019. Impact of ocean acidification on growth, onset of competence, and perception of cues for metamorphosis in larvae of the slippershell snail, Crepidula fornicata”.  Marine Biology 166:128.  https://doi.org/10.1007/s00227-019-3576-3


  • Andrade-Villagrán, P.V., K.S. Baria, J.A. Montory, J.A. Pechenik, and O.R. Chaparro. 2018. The influence of encapsulated embryos on the timing of hatching in the brooding gastropod Crepipatella dilatata. Journal of Sea Research 131: 69-78. (pdf)
  • Salas-Yanquin L, J.M. Navarro JM, J.A. Pechenik, J.A. Montory, and O.R. Chaparro. 2018. Volcanic ash in the water column: physiological impact on the suspension-feeding bivalve Mytilus chilensi. Marine Pollution Bulletin 127: 342-351.
  • Pechenik, J.A. 2018. Latent effects: surprising consequences of embryonic and larval experience on life after metamorphosis. In: Evolutionary Ecology of Marine Invertebrate Larvae. Oxford Univ. Press. T. Carrier, A. Reitzel, and A. Heyland, eds., pp 208-225. (pdf)
  • Andrade-Villagrán, P.V., D.A. Mardones-Toledo, F.J. Paredes-Molina, L.P. Salas-Tanquin, J.A. Pechenik, M. Matthews-Cascon, and O.R. Chaparro. Possible mechanisms of hatching from egg capsules in the gastropods Crepipatella dilatata and Crepipatella peruviana, species with different modes of early development.  Biological Bulletin 234:  69–84.
  • Kriefall, N.G., Pechenik, J.A., Pires, A, and S.W. Davies. 2018. Resilience of Atlantic slippersnail Crepidula fornicata larvae in the face of severe coastal acidification. Frontiers Mar. Sci. 5:312.      doi: 10.3389/fmars.2018.00312
  • Chaparro, O.R., L. P. Salas‑Yanquin, A. S. Matos, J. A. Bűchner‑Miranda,  M. W. Gray,  V. M. Cubillos, and J. A. Pechenik.  Pre‑hatching development in the intertidal zone negatively affects juvenile survival and physiology in the muricid gastropod Acanthina monodon. Marine Biology 165:155


  • Bashevkin, S.M., O.R. Chaparro, D. A. Mardones-Toledo, V.M. Cubillos, and J.A. Pechenik. 2017. Growing safe: acute size escape from desiccation in juvenile Crepipatella peruviana (Mollusca: Gastropoda). Biological Bulletin 233: 123-134. (pdf)
  • Pechenik. J.A., C.M. Diederich, O.R. Chaparro, J.A. Montory & A. Franklin. Differences in resource allocation to reproduction across the intertidal-subtidal gradient for two suspension-feeding marine gastropods: Crepidula fornicata and Crepipatella peruviana. Marine Ecology Progress Series 572: 165-178. (pdf)
  • Pechenik J.A. & R.R. Strathmann. The effect of age at metamorphosis on the transition from larval to adult suspension feeding of the slipper limpet Crepidula fornicata. Invertebrate Biology 136: 159-170. doi: 10.1111/ivb.12165 (pdf)
  • Burns, R.A. & J.A. Pechenik. The transcriptomic basis of metamorphic competence in the salt-marsh dwelling polychaete, Capitella teleta. Biological Bulletin 232: 158-170. (pdf)
  • Pechenik, J.A., Diederich, C.M., Browman, H.I., & J. Anders. Fecundity of the invasive marine gastropod Crepidula fornicata at the current northern extreme of its range. Invertebrate Biology136: 394-402. (pdf)


  • Montory, J.A., O.R. Chaparro, J.M. Navarro, J.A. Pechenik, and V.M. Cubillos. Post-metamorphic impact of brief hyposaline stress on recently hatched veligers of the gastropod Crepipatella peruviana (Calyptraeidae). Mar. Biol. 163:7 doi 10.1007/s00227-015-2791-9 (pdf)
  • Segura, C.J., J. A. Pechenik, J. A. Montory, J. M. Navarro, K. A. Paschke, V. M. Cubillos, and O. R. Chaparro. The cost of brooding in an estuary: Implications for gastropod females and their brooded embryos in the face of declining salinity. Marine Ecology Progress Series. Vol. 543: 187–199. doi: 10.3354/meps11593 (pdf)
  • Pechenik, J.A., O. Chaparro, A. Pilnick, M. Karp, M. Acquafredda, and R. Burns. Effects of embryonic exposure to salinity stress or hypoxia on post-metamorphic growth and survival of the polychaete Capitella teleta. Biological Bulletin 231: 103-112. (pdf)


  • Mardones-Toledo D.A., Montory J.A., Joyce A., Thompson R.J., Diederich C.M., Pechenik J.A., Mardones M.L., and O.R. Chaparro. Brooding in the Chilean oyster Ostrea chilensis: Unexpected complexity in the movements of brooded offspring within the mantle cavity. PLoS One 10(4): e0122859. doi:10.1371/journal.pone.0122859
  • Segura CJ, Montory JA, Cubillos VM, Diederich CM, Pechenik JA, Chaparro OR. Brooding strategy in fluctuating salinity environments: oxygen availability in the pallial cavity and metabolic stress in females and offspring in the Chilean oyster Ostrea chilensis. J. Comparative Physiology B 185: 659-68.
  • Pechenik, J. A, C. M. Diederich, O. R. Chaparro, V. M. Cubillos, and D. A. Mardones-Toledo. 2015. Role of the substrate in feeding and growth of the marine suspension-feeding gastropods Crepidula fornicata and Crepipatella peruviana. Biological Bulletin.
    Video 1: C peruviana cord ingestion with substrate
    Video 2: C peruviana losing cord, without substrate
    Video 3: C peruviana ingesting cord, on mesh
  • Pechenik, J.A., and A. S. Tyrell. Larval diet alters larval growth rates and post-metamorphic performance in the marine gastropod Crepidula fornicata. Marine Biology 162: 1597-1610. (pdf)
  • Pechenik, J.A., C. M. Diederich, and R. Burns. 2015. Shifts in hermit crab shell quality as a possible consequence of warmer New England winters. Marine Ecology Progress Series 529: 171–183 (pdf)
  • Pechenik, J.A., Casey M. Diederich, Robert Burns, Francesco Q. Pancheri, Luis Dorfmann. Influence of the commensal gastropod Crepidula plana on shell choice by the marine hermit crab Pagurus longicarpus, with an assessment of the degree of stress caused by different eviction techniques. J. Exp. Mar. Biol. Ecol. 469: 18–26. (pdf)
  • Bashevkin, S., and J.A. Pechenik. The interactive influence of temperature and salinity on larval and juvenile growth in the gastropod Crepidula fornicata (L.). J. Experimental Marine Biology and Ecology 470: 78-91. (pdf)
  • Diederich, C.M., O. R. Chaparro, D.A. Mardones-Toledo, G. P. Garrido, J.A. Montory, and J.A. Pechenik. Differences in feeding adaptations in intertidal and subtidal suspension-feeding gastropods: studies on Crepidula fornicata and Crepipatella peruviana. Marine Biology 162:1047–1059. (pdf)


  • Burns, R., J.A. Pechenik, W.J. Biggers, G. Scavo, and C. Lehman. 2014. The B vitamins nicotinamide (B3) and riboflavin (B2) stimulate metamorphosis in larvae of the deposit-feeding polychaete Capitella teleta: implications for a sensory ligand-gated ion channel. PLOS ONE 9 (Issue 11): | e109535 DOI: 10.1371/journal.pone.0109535 (pdf)
  • Montory, J.A., O.R. Chaparro, J.A. Pechenik, C.M. Diederich, V.M. Cubillos. 2014. Impact of short-term salinity stress on larval development of the marine gastropod Crepipatella fecunda (Calyptraeidae). J Experimental Marine Biology and Ecology 458: 39–45.
  • Chaparro, O.R., C. J. Segura, S. J. A. Osores, J. A. Pechenik, L. M. Pardo, and V. M. Cubillos. 2014. Consequences of maternal isolation from salinity stress for brooded embryos and future juveniles in the estuarine direct developing gastropod Crepipatella dilatata. Marine Biology 161:619–629. DOI 10.1007/s00227-013-2364-8 (pdf)
  • Montory, J.A., O. R. Chaparro, J. A. Pechenik, and C. Diederich. 2014. Effects of low salinity on adult behavior and larval performance in the intertidal gastropod Crepipatella fecunda (Calyptraeidae). PLoS ONE 9(7): e103820. doi:10.1371/journal.pone.0103820 (pdf)


  • Diederich, C.M., and J.A. Pechenik. 2013. Thermal tolerance of different Crepidula fornicata (Gastropoda) life history stages from intertidal and subtidal populations. Marine Ecology Progress Series. 486: 173–187. (pdf)
  • Mardones, M.L., O.R. Chaparro, J.A. Pechenik, C.J. Segura, and S.J.A. Osores. 2013. Embryonic brooding in Crepipatella fecunda: Implications for processes related to maternal feeding. J Experimental Marine Biology and Ecology 443: 141–146.
  • Le Cam, S,, F. Riquet, J. A. Pechenik, and F. Viard. 2013. Paternity and gregariousness in the sex-changing sessile marine gastropod Crepidula convexa: comparison with other protandrous Crepidula species. J. Heredity doi:10.1093/jhered/esu002


  • Pechenik, J.A., B. Fried and J. Bolstridge. 2012. The marine gastropods Crepidula plana and C. convexa do not serve as first intermediate hosts for larval trematode development. Comparative Parasitology 79: 5-8. (pdf)
  • Zelaya, D.G., J. A. Pechenik, C. S. Gallardo. (in press). Crepipatella dilatata (LAMARCK, 1822): An example of reproductive variability among gastropods. Nautilus.
  • Biggers, W.J. A. Pires, J. A. Pechenik, E. Johns, P. Priyam, T. Polson, and J. Polson. Inhibitors of nitric oxide synthase induce larval settlement and metamorphosis of the polychaete annelid Capitella teleta. Invertebrate Reproduction and Development 56: 1–13. (pdf)
  • Chaparro, O.R., L.A. Lincoqueo, A.J. Schmidt, D. Veliz & J.A. Pechenik. Comparing biochemical changes and energetic costs in gastropods with different developmental modes: Crepipatella dilatata and C. fecunda. Mar. Biol. 159:45–56.
  • Chaparro, O.R., L.A. Lincoqueo, A.J. Schmidt, D. Veliz & J.A. Pechenik. (in press). Oxygen constraints in the brood chamber: Anaerobic metabolism and the payment of oxygen debt in females and in brooded embryos of the limpet Crepipatella dilatata. Biol Bull.


  • O.R. Chaparro, J.A. Montory, J.A. Pechenik, V.M Cubillos, J.M. Navarro, and S.J. Osores. 2011. Ammonia accumulation in the brood chamber of the estuarine gastropod Crepipatella dilatata: How big a problem for mothers and brooded embryos? Marine Biology 410: 29–38.
  • C.M. Diederich, J.N. Jarrett, O.R. Chaparro, C.J. Segura, S.M. Arellano, J.A.Pechenik. 2011. Low salinity stress experienced by larvae does not affect post-metamorphic growth or survival in three calyptraeid gastropods. JEMBE 397: 94-105. (pdf)


  • Allen, J.D. and J.A. Pechenik. 2010. Understanding the Effects of Low Salinity on Fertilization Success and Early Development in the Sand Dollar Echinarachnius parma. Biol. Bull 218: 189-199. (pdf)
  • Segura, C.J., O.R. Chaparro, K.A. Paschke, J.A. Pechenik. 2010. Capsule walls as barriers to oxygen availability: Implications for the development of brooded embryos by the estuarine gastropod Crepipatella dilatata (Calyptraeidae). JEMBE 390: 49-57.
  • Pechenik, J.A., O.V. Ambrogio, and S. Untersee. 2010. Predation on juveniles of Crepidula fornicata by two crustaceans and two gastropods. J. Exp. Mar. Biol. Ecol. 384: 91-98. (pdf)


  • Montory, J.A., O.R. Chaparro, V.M. Cubillos, and J.A. Pechenik. 2009. Isolation of the incubatory cavity: alteration of pH and its impact on the protoconch of brooded veligers in the estuarine gastropod Crepipatella dilatata Lamarck (Gastropoda: Calyptraeidae). Mar. Ecol. Progr. Ser. 374: 157-166.
  • Chaparro, O.R., C.J. Segura, J.A. Montory, J. M. Navarro, and J.A. Pechenik. 2009. Brood chamber isolation during salinity stress in two estuarine mollusk species: from a protective nursery to a dangerous prison. Mar. Ecol Progr. Ser.374: 145-155. (pdf)
  • Le Cam, Sabrina, Jan A. Pechenik, Mathilde Cagnon, and Frédérique Viard. 2009. The influence of multiple paternity on larval growth rate variation in an invasive marine mollusk. J. Heredity 100: 455-464. (pdf)
  • Chaparro, O., J.A. Montory, C.J. Seguraand, and J. A. Pechenik. 2009. Effect of reduced pH on shells of brooded veligers in the estuarine bivalve Ostrea chilensis Philippi 1845. J. Exp. Mar. Biol. Ecol. 377: 107-112.
  • Ambrogio, O.V. and J.A. Pechenik. 2009. Do sex-changing male snails use mate choice to get a jump on their "size advantage"? Mar. Biol. 156:2173–2180. (pdf)


  • Thiyagarajan, V., J.A. Pechenik, L.A. Gosselin, and P.Y. Qian. 2007.
    Juvenile growth in barnacles: combined effect of delayed metamorphosis and sublethal exposure of cyprids to low salinity stress. Mar. Ecol.
    Progr. Ser. 344: 173-184.
  • Chiu, J.M, Thiyagarajan, V., Pechenik, J.A., and P.-Y. Qian. Environmental factors affect the composition of microbial films, subsequently influencing larval metamorphic behavior of the prosobranch gastropod Crepidula onyx. Marine. Biol. 151: 1417-1431.
  • Pechenik, J.A., and S. H. Levine. A new approach to estimating the magnitude of planktonic larval mortality using the marine gastropods Crepidula fornicata and C. plana. Marine Ecology Progress Series. 344: 107-118. (pdf)
  • Pechenik, J., J.S. Pearse, and P.-Y. Qian. Effects of salinity on spawning and early development of the tube-building polychaete Hydroides elegans in Hong Kong: not just the sperm's fault. Biological Bulletin. 212: 151-160. (pdf)
  • Pechenik, J.A., D.E. Cochrane, W.Li, E.T.West, A.Pires, M.Leppo. Nitric oxide inhibits metamorphosis in larvae of Crepidula fornicata, the slippershell snail. Biol. Bull. 213. 160-171. (pdf)
  • Li, W. and J. A. Pechenik.  The impact of inbreeding on reproduction and juvenile performance in two marine gastropods with contrasting reproductive patterns.  Marine Ecology Progress Series 346: 219–234. (pdf)
  • Untersee, S. and J. A. Pechenik.  Local adaptation and maternal effects in two species of marine gastropod (genus Crepidula) that differ in dispersal potential.  Marine Ecology Progress Series 347: 79-85. (pdf)


  • Pechenik, J.A. Larval experience and latent effects--metamorphosis is not a new beginning.  Integrative and Comparative Biology.  J. Integr. Comp. Biol. 47: 1-11.  (pdf)


  • Li, W., and J. A. Pechenik, 2004. A forced association between the slippersnail Crepidula convexa and the hermit crab Pagurus longicarpus?: possible control by a third party. J. Exp. Mar. Biol. Ecol. 311: 339-354 (pdf)
  • Gravel, B.E., Wong, P.E., Starks, P.T., and J.A. Pechenik. 2004. The use of artificial shells for exploring shell preference in the marine hermit crab Pagurus longicarpus (Say). Acta Zool. Fennica 41: 477-485.
  • Pechenik, J.A., M. Blanchard, and R. Rotjan.2004. Susceptibility of larval Crepidula fornicata to predation by suspension-feeding adults. J. Exp. Mar. Biol. Ecol. 306: 75-94.


  • Marshall, D.J., Pechenik, J.A., and M.J. Keough. Larval activity levels and delayed metamorphosis affect post-larval performance in the colonial ascidian Diplosoma listerianum. Mar. Ecol. Progr. Ser. 246: 153-162.
  • Pechenik, J.A., Marsden, I.D., and O. Pechenik. Effects of temperature, salinity, and air exposure on development of the estuarine pulmonate gastropod Amphibola crenata. J. Exp. Mar. Biol. Ecol. 292: 159-176.


  • Pechenik, J.A., Wei Li, and David E. Cochrane. Timing is everything: The effects of putative dopamine antagonists on metamorphosis vary with larval age and experimental duration in the prosobranch gastropod Crepidula fornicata Biological Bulletin 202: 137-147.
  • Pechenik, J.A., J. Jarrett, and J. Rooney. Relationship between larval nutritional experience, larval growth rates, and juvenile growth rates in the prosobranch gastropod Crepidula fornicata. J. Exp. Mar. Biol. Ecol. 280: 63-78.


  • Pechenik, J.A., and M.E. Rice. Influence of delayed metamorphosis on post settlement survival and growth of the sipunculan Apionsoma misakiana. Invert. Biol. 120: 50-57.
  • Pechenik, J.A., Berard, R., Daniels, D., Gleason, T.R., and D. Champlin. Influence of lowered salinity and elevated cadmium on the survival and metamorphosis of Capitella sp. I trochophores. Invert. Biol. 120: 142-148.
  • Pechenik, J.A., J.M. Reed, and M. Russ. Should auld acquaintance be forgot: Possible influence of computer databases on citation patterns in the biological literature. BioScience 51: 583-588.
  • Pechenik, J.A. Life Cycles. In: C.W. Fox, D. Roff, and F. Fairbairn, eds., Evolutionary Ecology: Perspectives and Case Studies, Oxford University Press, NY.
  • Pechenik, J.A. and B. Fried. Crepidula fornicata is not a first intermediate host for trematodes: Who is? J. Exp. Mar. Biol. Ecol. 261: 211-224.
  • Pechenik, J.A., J. Hsieh, S. Owara, S. Untersee, D. Marshall, and W. Li. Factors selecting for avoidance of drilled shells by the hermit crab Pagurus longicarpus. J. Exp. Mar. Biol. Ecol. 262: 75-89.
  • Pechenik, J.A., T. Gleason, D. Daniels, and D. Champlin. Influence of larval exposure to salinity and cadmium stress on juvenile performance of two marine invertebrates (Capitella sp. I and Crepidula fornicata). J. Exp. Mar. Biol. Ecol. 264: 101-114.
  • Gaudette, M.F., J. Lowther, and J. A. Pechenik. Heat shock induces metamorphosis in larvae of the prosobranch gastropod Crepidula fornicata. J. Exp. Mar. Biol. Ecol. 266: 151-164.


  • Pechenik, J.A., Berard, R., and L. Kerr. Effects of reduced salinity on survival, growth, reproductive success, and energetics of the euryhaline polychaete Capitella sp. I. J. Exp. Mar. Biol. Ecol. 254: 19-35.
  • Klinzing, M.S., and J.A. Pechenik. Evaluating whether velar-lobe size indicates food limitation among larvae of the marine gastropod Crepidula fornicata. J. Exp. Mar. Biol. Ecol. 252: 255-279.
  • Pechenik, J.A., Lewis, S. Avoidance of drilled gastropod shells by the hermit crab Pagurus longicarpus at Nahant, Massachusetts. J. Exp. Mar. Biol. Ecol. 253:17-32.


  • Pechenik, J.A. On the benefits of being a larva... or not. In Aquatic Life Cycle Strategies, M. Whitfield, J. Matthews, C. Reynolds (eds.), pp. 97-104. Marine Biological Association of the United Kingdom, Plymouth.
  • Pechenik, J.A. Reproductive Modes of Marine Invertebrates. In, Encyclopedia of Reproduction, Academic Press, NY. Volume 3: 98-103.
  • Fried, B., L.S. Eyster, and J.A. Pechenik. Histochemical glycogen and neutral lipid in Echinostoma trivolvis cercariae and effects of exogenous glucose on cercarial longevity. J. Helminthol. 72: 83-85.
  • Pechenik, J.A. On the advantages and disadvantages of larval stages in benthic marine invertebrate life cycles. Marine Ecol. Progr. Series. 177: 269-297.
  • Cohen, R.,and J.A. Pechenik. Relationship between sediment organic content, metamorphosis, and postlarval performance in the deposit-feeding polychaete Capitella sp. I. J. Exp. Marine Biol. Ecol. 240:1-18. (pdf)
  • Hilbish, T.J., Sasada, K., Eyster, L.S., and J.A. Pechenik. Relationship between rates of swimming and growth in veliger larvae: genetic variance and covariance. J. Exp. Marine Biol. Ecol. 239:183-193.
  • Pechenik, J.A. Metamorphosis. Encyclopedia of Science and Technology, McGraw Hill, NY.


  • Pechenik, J.A., D. Wendt, and J. Jarrett. Metamorphosis is not a new beginning: Larval experience reduces rates of postlarval growth, development, and survival in marine invertebrates. BioScience. 48:901-910. (pdf)
  • Pechenik, J.A., and P. Y. Qian. Onset and maintenance of metamorphic competence in the marine polychaete Hydroides elegans in response to three chemical cues. J. Exp. Marine. Biol. Ecol. 226:51-74.
  • Qian, P. Y., and J.A. Pechenik. Effects of starvation and delayed metamorphosis on juvenile survival and growth of the tube dwelling polychaete Hydroides elegans. J. Exp. Marine. Biol. Ecol. 227:169-185. (pdf)


  • Jarrett, J.N., and J.A. Pechenik. Temporal variation in cyprid quality and juvenile growth capacity for the barnacle Semibalanus balanoides. Ecology 78:1262-1265. (pdf)
  • Pechenik, J.A. Getting the most out of student laboratory reports. Strategies 23:4-5.


  • Pechenik, J.A., T.J. Hilbish, L.S. Eyster, and D. Marshall. Relationship between larval and juvenile growth rates in two marine gastropods, Crepidula plana and C. fornicata. Marine Biol. 125:119-127. (pdf)
  • Pechenik, J.A., K. Hammer, and C. Weise. The effect of starvation on acquisition of competence and postmetamorphic performance in the marine prosobranch gastropod Crepidula fornicata (L.). J. Exp. Mar. Biol. Ecol. 199:137-152. (pdf)
  • Pechenik, J.A., S. Estrella, and K. Hammer. Food limitation stimulates metamorphosis and alters postmetamorphic growth rate in the marine prosobranch gastropod Crepidula fornicata. Marine Biol. 127:267-275. (pdf)


  • Pechenik, J.A., Hadfield, M.G., and L.S. Eyster. Assessing whether larvae of the opisthobranch gastropod Phestilla sibogae become responsive to three chemical cues at the same age. J. Exp. Mar. Biol. Ecol. 191:1-17.
  • Pechenik, J.A., and B. Fried. Effect of temperature on survival and infectivity of Echinostoma trivolvis cercariae: a test of the energy limitation hypothesis. Parasitology 111:373-378.


  • Pechenik, J.A., D. Rittschof, and A.R. Schmidt. Influence of delayed metamorphosis on survival and growth of juvenile barnacles Balanus amphitrite. Marine Biol. 115: 287-294.  (pdf)
  • Pechenik, J.A., and C.C. Gee. Onset of metamorphic competence in larvae of the gastropod Crepidula fornicata, judged by a natural and an artificial cue. J. Exp. Mar. Biol. Ecol. 167: 59-72.


  • Pechenik, J.A., and J.S. Tashiro. The Graphing Detective: An exercise in critical reading, experimental design & data analysis. Amer. Biol. Teacher. 54: 432-435.  (pdf)
  • Rice, M.A., and J.A. Pechenik. A review of the factors influencing the growth of the northern quahog, Mercenaria mercenaria (Linnaeus, 1758). J. Shellf. Res. 11: 279-287.


  • Pechenik, J.A. and J. Shiro Tashiro. Instant animals and conceptual loops: Teaching experimental design, data analysis, and scientific writing. Amer. Biol. Teacher 53(4): 220-228. (pdf)
  • Zimmerman, K.M., and J.A. Pechenik. How do temperature and salinity affect relative rates of growth, morphological differentiation, and time to metamorphic competence in larvae of the marine gastropod Crepidula plana? Biol. Bull. 180: 372-386.
  • Pechenik, J., and T. Cerulli. Influence of delayed metamorphosis on survival, growth, and reproduction of the marine polychaete Capitella sp. I. J. Exp. Marine Biol. Ecol. 151: 17-27.


  • Pechenik, J.A., L.S. Eyster, J. Widdows, and B.L. Bayne. The influence of food concentration and temperature on growth and morphological differentiation of blue mussel larvae, Mytilus edulis L. J. Exp. Marine Biol. Ecol. 136: 47-64. (pdf)
  • Pechenik, J.A. Delayed metamorphosis by larvae of benthic marine invertebrates: Does it occur? Is there a price to pay? Ophelia 32: 63-94.


  • Woollacott, R.M., J.A. Pechenik, and K.M. McSorley. Influence of swimming time on early colony development in Bugula stolonifera (Bryozoa: Cheilostomata). Marine Biol. 102: 57-63.
  • Pechenik, J.A., and L.S. Eyster. Influence of delayed metamorphosis on the growth and metabolism of young Crepidula fornicata (Gastropoda) juveniles. Biological Bulletin 176: 14-24.


  • Pechenik, J.A. and L.S. Eyster. Comparison of growth, respiration, and feeding of juvenile Crepidula fornicata (L.) following natural or KCl-triggered metamorphosis. J. Exp. Marine Biol. Ecol. 118: 269-279.


  • Dobberteen, R.A., and Pechenik, J.A. Comparison of larval bioenergetics for two marine gastropods with widely differing lengths of planktonic life (Thais haemastoma canaliculata (Gray) and Crepidula fornicata (L.)). J. Exp. Mar. Biol. Ecol. 109: 173-191.
  • Pechenik, J.A. Environmental influences on larval survival and growth. IN, A.C. Giese and J.S. Pearse (eds.). Reproduction of Marine Invertebrates, Vol. 9, Blackwell Scientific, NY., pp. 551-608.
  • Pechenik, J.A., and W.D. Heyman. The influence of KCl and other chemicals on metamorphosis of the marine gastropod Crepidula fornicata (L.). J. Exp. Mar. Biol. Ecol. 112: 27-38.
  • Eyster, L.S., and J.A. Pechenik. Attachment of Mytilis edulis L. larvae on algal and byssal filaments is enhanced by water agitation. J. Exp. Mar. Biol. Ecol. 114: 99-110.


  • Pechenik, J.A. Field evidence for delayed metamorphosis of larval gastropods: Crepidula plana Say, C. fornicata (L.), and Bittium alternatum Say. J. Exp. Mar. Biol. Ecol. 97: 313-319. (pdf)
  • Greenberger, J.S., Pechenik, J.A., Lord, L., Gould, L., Naparstek, E., Kase, K., and T.J. FitzGerald. X-irradiation effects on growth and metamorphosis of gastropod larvae (Crepidula fornicata): a model for environmental radiation teratogenesis. Archives of Environ. Contam. Toxicol. 15: 227-234.
  • Turner, R.D., Pechenik, J.A., and C.B. Calloway. The language of marine invertebrate developmental patterns--problems and needs. In: M.-F. Thompson, R. Sarojini, and R. Nagabhushanam, eds. Biology of Benthic Marine Invertebrates, pp. 227-235. Oxford and IBH Publ. Co., New Delhi.
  • Pechenik, J.A. The encapsulation of eggs and embryos by molluscs: an overview. Amer. Malac. Bull. 4: 165-172.


  • Pechenik, J.A. Delayed metamorphosis of marine molluscan larvae: current status and directions for future research. Amer. Malac. Bull. Special Ed. No. 1: 85-91.
  • Lima, G.M., and Pechenik, J.A. The influence of temperature on growth rate and length of larval life of the gastropod Crepidula plana Say. J. Exp. Mar. Biol. Ecol. 90: 55-71.


  • Pechenik, J.A. Relationship between rate of development and duration of larval life in larvae of the marine prosobranch gastropod Crepidula fornicata. J. Exp. Mar. Biol. Ecol. 74: 241-257.
  • Pechenik, J.A., S.C. Chang, and A. Lord. Studies on the encapsulated development of the marine gastropod, Nucella lapillus. Marine Biol. 78: 223-229.
  • Pechenik, J.A. Influence of temperature and temperature shifts on the development of the chiton, Mopalia muscosa. Int. J. Invert. Reprod. Devel. 7: 3-12.
  • Pechenik, J.A., and G. Lima. Relationship between growth, differentiation, and duration of larval life in individually-reared larvae of the marine gastropod Crepidula fornicata. Biol. Bull. 166: 537-549. (pdf)
  • Pechenik, J.A., R.S. Scheltema, and L.S. Eyster. Growth-stasis and reduced shell calcification in larvae of the gastropod Cymatium parthenopeum during trans-Atlantic transport. Science, 224: 1097-1099.
  • Pechenik, J.A. Predicting the consequences of water-soluble pollutant input into the marine environment. Hazardous Waste, 1: 273-282.


  • Pechenik, J.A. Egg capsules of Nucella lapillus protect against low-salinity stress. J. Exp. Marine Biol. Ecol. 71: 165-179. (pdf)
  • Pechenik, J.A., and D.M. Miller. Increased susceptibility to No. 2 fuel oil coincident with the initiation of particle feeding in developing mud snails, Ilyanassa obsoleta. Estuaries 6: 237-242.


  • Pechenik, J.A. Ability of some gastropod egg capsules to protect against low-salinity stress. J. Exp. Marine Biol. Ecol. 63: 195-208. (pdf)


  • Pechenik, J.A. Growth and energy balance during the larval life of three prosobranch gastropods. J. Exp. Mar. Biol. Ecol. 44: 1-28. (pdf)
  • Johns, D.M., and J.A. Pechenik. Effect of exposure to No. 2 fuel oil on growth and energetics of rock crab larvae, Cancer irroratus. Marine Biol. 55: 247-254.


  • Pechenik, J.A. Leakage of ingested carbon by gastropod larvae, and its effect on the calculation of assimilation efficiency. Estuaries 2: 45-49.
  • Pechenik, J.A. The role of encapsulation in the life histories of marine invertebrates. Amer. Naturalist 114: 859-870. (pdf)
  • Pechenik, J.A., F. Perron, and R.D. Turner. The role of phytoplankton in the diets of adult and larval shipworms, Lyrodus pedicellatus (Bivalvia, Teredinidae). Estuaries 2: 58-60.
  • Pechenik, J.A., and N.S. Fisher. Ingestion and assimilation of three phytoplankton species related to larval growth rates of the mud snail, Nassarius obsoletus. J. Exp. Mar. Biol. Ecol. 38: 57-80.
  • Pechenik, J.A., D.M. Johns, and D.C. Miller. Effects of exposure to No. 2 fuel oil on the survival and reproductive biology of some marine invertebrates. Proc. Symposium on the State of Marine Environmental Research, June 15-16, 1977. Environmental Research Laboratory, USEPA, Narragansett, R.I., pp. 135-156.


  • Pechenik, J.A. Adaptations to intertidal development: Studies on Nassarius obsoletus. Biol. Bull. 154: 282-291.
  • Pechenik, J.A. Winter reproduction in the gastropod Nassarius trivittatus. Veliger 21: 297-298.


  • Sastry, A.N., and J.A. Pechenik. A review of the ecology, physiology and behavior of lobster larvae (Homarus americanus and H. gammarus). Div. Fish. Oceanogr. Circ. 7, Commonwealth Scientific and Industrial Research Org., Australia, pp. 159-173.


  • Pechenik, J.A. The escape of veligers from the egg capsules of Nassarius obsoletus and Nassarius trivittatus (Gastropoda, Prosobranchia). Biol. Bull. 149: 580-589. (pdf)